C. Casiot, G. Morin, F. Juillot, O. Bruneel, J. Personné et al., , p.408

V. Bonnefoy and F. Elbaz-poulichet, Bacterial Immobilization and Oxidation of Arsenic in Acid 409

, Mine Drainage (Carnoulès Creek, France), Wat Res, vol.37, pp.2929-2965, 2003.

F. Battaglia-brunet, Y. Itard, F. Garrido, F. Delorme, C. Crouzet et al., A 411 simple biogeochemical process removing arsenic from a mine drainage water, Geomicrobiol, vol.412, pp.201-211, 2006.

C. C. Fuller, J. A. Davis, and G. A. Waychunas, Surface chemistry of ferrihydrite: Part 2. Kinetics 414 of arsenate adsorption and coprecipitation, Geochim Cosmochim Acta, pp.227-415, 1993.

P. L. Smedley and D. G. Kinniburgh, A review of the source, behaviour and distribution of 417 arsenic in natural waters, Appl Geochem, vol.17, pp.517-568, 2002.

L. C. Anderson and K. W. Bruland, Biogeochemistry of Arsenic in Natural Waters: The 444 Importance of Methylated Species, Environ Sci Technol, vol.25, pp.420-427, 1991.

D. Muller, D. Lievremont, D. D. Simeonova, J. C. Hubert, and M. C. Lett, , p.446

, Genes from a Metal-Resistant-Proteobacterium, J Bacteriol, vol.185, pp.135-141, 2003.

Q. Wang, Y. Han, and K. Shi, An Oxidoreductase AioE is Responsible for Bacterial 448

, Arsenite Oxidation and Resistance. Scientific Reports, vol.7, p.41536, 2017.

J. M. Santini, R. N. Van-den-hoven, and . Molybdenum, Containing Arsenite Oxidase of the, p.450

, Chemolithoautotrophic Arsenite Oxidizer NT-26, J Bacteriol, vol.186, pp.1614-1619, 2004.

R. S. Oremland, S. E. Hoeft, J. M. Santini, N. Bano, R. A. Hollibaugh et al., Anaerobic 453 Oxidation of Arsenite in Mono Lake Water and by a Facultative, Arsenite-Oxidizing, p.454

. Chemoautotroph, Strain MLHE-1. Appl Environ Microbiol, vol.68, pp.4795-4802, 2002.

J. M. Santini, L. I. Sly, A. Wen, D. Comrie, P. D. Wulf-durand et al., , p.456

, Oxidizing Bacteria Isolated from Australian Gold Mining Environments-Phylogenetic 457

, Relationships. Geomicrobiol J, vol.19, pp.67-76, 2002.

F. Battaglia-brunet, E. Achbouni, H. Quemeneur, M. Hallberg, K. B. Kelly et al.,

, Proposal that the arsenite-oxidizing organisms, Thiomonas cuprina and 'Thiomonas 460 arsenivorans' be reclassified as strains of Thiomonas delicata, p.461

, Thiomonas delicata. Intl J Syst Environ Microbiol, vol.61, pp.2816-2821, 2011.

S. Challan-belval, F. Garnier, C. Michel, S. Chautard, D. Breeze et al., Enhancing 463 pozzolana colonization by As(III)-oxidizing bacteria for bioremediation purposes, Appl, vol.464

, Microbiol Biotechnol, vol.84, pp.565-573, 2009.

S. P. Bachate, R. M. Khapare, and K. M. Kodam, Oxidation of arsenite by two beta-466 proteobacteria isolated from soil, Applied Microbiology and Biotechnology, vol.93, pp.2135-2145, 2012.

T. Lescure, J. Moreau, C. Charles, B. A. Saanda, T. Thouin et al.,

I. Battaglia-brunet and F. , Influence of organic matters on AsIII oxidation by the microflora of 469 polluted soils, Env Geochem Health, vol.38, pp.911-925, 2016.

V. S. Nandre, S. P. Bachate, and R. C. Salunkhe, Enhanced Detoxification of Arsenic Under 471 Carbon Starvation: A New Insight into Microbial Arsenic Physiology, Curr Microbiol, vol.472, p.614, 2017.

A. M. Spuches, H. G. Kruszyna, A. M. Rich, and D. E. Wilcox, Thermodynamics of the AsIII-thiol 474 interaction: arsenite and monomethylarsenite complexes with glutathione, dihydroglucoic acid 475 and other thiol ligants, Inorg Chem, vol.44, pp.2964-2972, 2005.

F. Battaglia-brunet, C. Joulian, D. Breeze, L. Poirier, S. Coulon et al., , p.477, 2010.

, Monitoring of arsenic bio-oxidation and bio-redution activities in highly As-polluted soils 478 during a phytostabilization field experiment (Phytoperf project), Consoil 2010, 11th 479 International UFZ-Deltares/TNO Conference on Management of Soil, Groundwater and 480 Sediment

D. Muller, D. D. Simeonova, P. Riegel, S. Mangenot, S. Koechler et al., , p.482

M. C. Lett, Herminiimonas arsenicoxydans sp. nov., a metalloresistant bacterium, Int J Syst, p.483
URL : https://hal.archives-ouvertes.fr/hal-00113703

, Evol Microbiol, vol.56, pp.1765-1769, 2003.

W. Weeger, D. Lièvremont, M. Perret, F. Lagarde, J. C. Hubert et al.,

, Oxidation of arsenite to arsenate by a bacterium isolated from an aquatic environment, Biometals, vol.12, pp.141-149, 1999.

F. Battaglia-brunet, M. C. Dictor, F. Garrido, C. Crouzet, D. Morin et al., , p.488

P. Baranger, An arsenic(III)-oxidizing bacterial population: selection, characterisation, and 489 performance in reactors, J Appl Microbiol, vol.93, pp.656-667, 2002.

M. Quéméneur, Les processus biogéochimiques impliqués dans la mobilité de l'arsenic : 491 recherche de bioindicateurs, 2008.

L. Fernandez-rojo, M. Hery, L. Pape, P. Braungardt, C. Desoeuvre et al., , p.495

E. Resongles, E. Laroche, S. Delpoux, C. Joulian, F. Battaglia-brunet et al., , p.496

G. Morin and C. Casiot, Biological attenuation of arsenic and iron in a continuous flow 497 bioreactor treating acid mine drainage (AMD), Water Research, vol.123, pp.594-606, 2017.

D. Slyemi and V. Bonnefoy, How prokaryotes deal with arsenic, Env Microbiol Reports, vol.499, pp.571-586, 2012.

A. Hovasse, O. Bruneel, C. Casiot, A. Desoeuvre, J. Farasin et al., , vol.501

C. Carapito, F. Arsène-ploetze, and . Spatio, Temporal Detection of the Thiomonas Population and 502 the Thiomonas Arsenite Oxidase Involved in Natural Arsenite Attenuation Processes in the 503

, Carnoulès Acid Mine Drainage. Front Cell Dev Biol, vol.4, pp.1-14, 2016.

A. Lieutaud, R. Van-lis, S. Duval, L. Capowiez, D. Muller et al., , p.505

L. Ml, . Mc, W. Nitschke, and B. Shoepp-cothenet, Arsenite oxidase from Ralstonia sp. 22: 506 characterization of the enzyme and its interaction with soluble cytochromes, Journal of 507 Biological Chemistry, vol.285, pp.2433-2441, 2010.

D. R. Kashyap, L. M. Botero, W. L. Franck, D. J. Hassett, and T. R. Mcdermott, Complex regulation 509 of arsenite oxidation in Agrobacterium tumefaciens, J Bacteriol, vol.188, pp.1081-1088, 2006.

J. Poirel, C. Joulian, C. Leyval, and P. Billard, Arsenite-induced changes in abundance and 511 expression of arsenite transporter and arsenite oxidase genes of a soil microbial community

, Res Microbiol, vol.164, pp.457-465, 2013.

S. Koechler, F. Arsène-ploetze, C. Brochier-armanet, G. , and F. ,

A. Salmeron, B. Jost, D. Lièvremont, M. Philipps, F. Plewniak et al.,

, Constitutive arsenite oxidase expression detected in arsenic-hypertolerant Pseudomonas 516 xanthomarina S11, Res Microbiol, vol.166, pp.205-214, 2015.

S. Koechler, J. Cleiss-arnold, C. Proux, O. Sismeiro, M. A. Dillies et al., , p.518

F. Hommais, D. Lievremont, F. Arsene-ploetze, J. Y. Coppee, and P. N. Bertin, Multiple controls affect 519 arsenite oxidase gene expression in Herminiimonas arsenicoxydans, BMC Microbiol, vol.520, p.53, 2010.

J. M. Janda and S. L. Abbott, 16S rRNA gene sequencing for bacterial identification in the 522 diagnostic laboratory: pluses, perils, and pitfalls, J Clin Microbiol, vol.45, pp.2761-2764, 2007.

T. M. Salmassi, K. Venkateswaran, M. Satomi, N. K. Newman, D. K. Hering et al.,

, Oxidation of arsenite by Agrobacterium albertimagni AOL15, sp. Nov., isolated from hot 525 creek California, Geomicrobiol J, vol.19, pp.53-66, 2002.

J. M. Santini, L. I. Sly, R. D. Schnagl, and J. M. Macy, A new chemolithoautotrophic arsenite-527 oxidizing bacterium isolated from a gold mine: phylogenetic, physiological, preliminary 528 biochemical studies, Appl Environ Microbiol, vol.66, pp.92-97, 2000.

P. Ghosh, B. Rathinasabapathi, M. Teplitski, and L. Q. Ma, Bacterial ability in AsIII oxidation 530 and AsV reduction: Relation to arsenic tolerance, P uptake, and siderophore production, Chemosphere, vol.531, issue.138, pp.995-1000, 2015.

K. T. Kitchin and K. Wallace, The role of protein binding of trivalent arsenicals in arsenic 533 carcinogenesis and toxicity, J Inorg Biochem, vol.102, pp.532-539, 2008.

A. R. Achour, P. Bauda, and P. Billard, Diversity of arsenite transporter genes from arsenic-535 resistant soil bacteria, Res Microbiol, vol.158, pp.128-137, 2007.

A. Achour-rokbani, A. Cordi, P. Poupin, P. Bauda, and P. Billar, Characterization of the ars 537 gene cluster from extremely arsenic-resistant Microbacterium sp. strain A33, Appl Environ 538 Microbiol, vol.76, pp.948-955, 2010.