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R. A. , R. A. Arc-1215551-to, R. A. , and A. K. , Postdoctoral Fellowship and by National Science Foundation grants EAR-1024929 to a LabEx International Research Chair funded by the 'Laboratoire d'Excellence' LabexMER(ANR-10-LABX-19) and co-funded by a grant from the French government under the programme, 'Investissements d'Avenir', and a funded visiting Associate Professor position at the Natural History Museum of Denmark. A.K.T. was supported by a Hellman Fellowship, and National Science Foundation grant