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P. Mein, Kachacha for his commitment We are indebted to I. Antoñanzas-Asso for sorting the sediment We sincerely thank C. Denys and C. Argot (Muséum national d'Histoire naturelle, Paris) and F. Mayer and S. Jancke (Museum für Naturkunde, Berlin) for making available to RLA the ctenodactylid material under their care. L. Marivaux and B. Marandat (Université Montpellier 2, Montpellier) kindly answered our enquiries. We also express our gratitude to A, Acknowledgments We acknowledge very much the students and) for having improved the manuscript. M. Furió, A. García, C. Paradela, and L. Tormo (Museo nacional de Ciencias naturales-CSIC, Madrid) nicely took the ? CT scan images and SEMs